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It has also been indicated that riboflavin deficiency decreases the mRNA level of antioxidant enzymes in muscle (GR, GPx, SOD and CAT) (Reference Jiang, Chen and Liu33) , intestine (GR, GPx and SOD) (Reference Chen, Feng and Jiang34) and gills (GR, SOD) (Reference Chen, Feng and Jiang35) of fish